1876-1877

Welch journeys to Strasbourg to study pathological anatomy via laboratory courses with the microscope (Flexner, 78-79), then on to Leipzig, where he studied pathology with Ernst Wagner and physiology with Carl Ludwig (Flexner, 82-88), who gave him an Arbeit microscopic investigation of nerves and ganglion cells of the heart (85); then to Breslau, where he studies experimental pathology with Julius Cohnheim. Cohnheim’s Arbeit for Welch was edema of the lungs (which problem Welch solved. To wit, when left ventricle was paralyzed and right ventricle continued to pump blood to the lungs, edema resulted) (Flexner, 94-99).